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ALABAMA QUAIL PROJECT
2 YEAR PROGRESS REPORT

Field work for the Alabama Quail Project has been underway since the fall of 2002 on Creekstand and Sehoy Plantations. The main objectives were to determine the limiting factors of wild quail populations on these properties by conducting year-round telemetry monitoring of a large sample of quail. In addition to telemetry work, we have conducted quail covey and predator censuses on the sites each year. Over the course of the last 2 years, 400 wild quail have been radio-tagged on Sehoy and Creekstand. Our field work is broken down into 2 seasons: Fall through winter - the over-winter period, and spring through summer - the reproductive period. The radios have been evenly distributed between study sites and between 2 Fall/Winter and 2 Spring/Summer periods. These radio-tagged birds are beginning to provide us with insights into factors that are making these quail populations tick, such as data on adult survival, habitat use, and reproductive output. Following is a summary of our data to date.

Survival

Fall/Winter Survival
The graph in Figure 1 represents annual survival for the Alabama properties during 02/03 and 03/04 and compares these 2 years to the long term over-winter average in Georgia. Over-winter survival continues to be surprisingly good on the study sites. Our long term studies have shown that the number of birds surviving to the breeding season is one of the primary driving forces behind population performance. Good winter cover combined with supplemental feeding is apparently keeping the birds safe through the over-winter period.

Spring/Summer Survival
Spring/Summer survival has been monitored for 2 years. Backing up previous studies, we are finding that higher percentages of unburned cover equals higher quail survival, nest production, and nest success. During spring 2003, Site 1 had 10% of the quail cover unburned, and Site 2 had approximately 70% unburned. Spring/Summer survival was significantly higher on Site 2 (see Figure 2.). The amount of cover burned in spring 2004 was much different for these same areas. On Site 1, permanent fire lanes were established before burning, and unburned cover increased to 50%. Survival increased significantly over the previous spring/summer on Site 1 (Figure 3.). On Site 2 in 2004, the amount of unburned cover decreased to 10% and survival (Figure 4.) was much lower compared to Site 1 with 50% unburned. Clean burning over many acres is obviously a limiting factor to quail populations, because the birds are exposed to predation and available nest sites are reduced. Unburned cover should range between 30 and 50% of a quail hunting course.

 


Predation

The causes of death for radio-tagged quail on each study site and annual percentages for each predator type are represented in Tables 1 and 2. As shown by these tables, avian predators are responsible for a high percentage of over-winter mortality, while other predator types make up the lower percentages.

Table 1: Site 1 Causes of Adult Mortality
Predator Types Nov. 02 - Mar. 03 Apr. 03 - Sept. 03 Annual
02 - 03
Nov. 03 - Mar. 04 Apr. 04 - Sept. 04 Annual
03 - 04
Nov. 02 - Sept. 04
Avian 9 30 39 (61%) 7 26 33 (68%) 72 (64%)
Mammal 0 12 12 (19%) 1 5 6 (12%) 18 (16%)
Hunt 4 0 4 (6%) 2 0 2 (4%) 6 (5%)
Unknown 2 7 9 (14%) 3 4 7 (14%) 16 (14%)
Snake 0 0 0 0 1 1 (2%) 1 (1%)
Total 15 49 64 13 36 49 113

Table 2: Site 2 Causes of Adult Mortality
Predator Types Nov. 02 - Mar. 03 Apr. 03 - Sept. 03 Annual
02 - 03
Nov. 03 - Mar. 04 Apr. 04 - Sept. 04 Annual
03 - 04
Nov. 02 - Sept. 04
Avian 9 25 34 (59%) 7 21 28 (57%) 62 (58%)
Mammal 2 6 8 (14%) 1 12 13 (27%) 21 (20%)
Hunt 6 0 6 (10%) 4 0 4 (8%) 10 (9%)
Unknown 4 4 8 (14%) 1 2 3 (6%) 11 (10%)
Snake 0 2 2 (3%) 0 1 1 (2%) 3 (3%)
Total 21 37 58 13 36 49 107

For both study sites, predation significantly increases by all predator types, except human, during the nesting season. Again, this is after spring burning when birds are more exposed. Also, the coveys are broken up and don’t have safety in numbers and are more vulnerable to mammalian predation while nesting.

Besides data collected from radio-tagged quail, the predation issue continues to be examined by a scent station survey. This survey gives an index to the mammalian predator population on the study sites. The survey is conducted each June and October. A scented disk is placed in a 3 foot diameter circle of tracking sand every ½ mile along roads and checked like a trap line. Twenty stations were placed on each study site and checked for 5 consecutive nights. The index is the average number of “hits” per night by known nest predators (raccoons, opossums, armadillos, bobcats) calculated as a percentage. Table 3 represents the predator indexes over the past 2 years on the study sites.

Table 3: Study Site Predator Indexes
  June 2003
% Hit
October 2003
% Hit
June 2004
% Hit
October 2004
% Hit
SITE 1 13% 20% 8% 6%
SITE 2 13% 19% 17% 17%

An intensive predator control program was implemented on Site 1 following the October 2003 survey. As evidenced by Table 3, the mammalian predator index continues to decrease on Site1. The annual percent of mammalian predation on Site 1 has also decreased after initiation of the predator control program, as seen in Table 1.


Reproduction

One of the most important parameters we measure is reproductive output from radio-tagged quail. Table 4 represents reproductive parameters for each study site over the last 2 nesting periods. Past experience has shown that total reproductive performance is greatly affected by poor adult survival and a high mammalian predator count. As recorded in Table 4, the best overall reproduction numbers in 2003 were on Site 2, and in 2004 were on Site 1. These were nesting periods in which adequate cover remained after burning and mammalian predator counts were at their lowest recorded numbers.

  # hens alive
April 1
# nests total
(male)
Hatching Success % (early, late) # Broods
(male)
Depredated
(male)
Still Incubating Nests per 100 hens Broods per 100 hens (with males)
Site 1
2003
56 25
(2)
52%
67%, 14%
13 12
(2)
0 41.07 23.21
Site 2
2003
44 24 50%
50%, 50%
12 12 0 54.55 27.27
Site 1
2004
56 37
(3)
57%
52%, 63%
21
(2)
16
(1)
0 60.71 33.93
(37.50)

Site 2
2004

48 32
(3)
31
33%, 29%
10
(1)
22
(2)
0 60.42 18.75
(20.83)

Popular Trends
We have conducted a covey call population census for each study site during fall of 2002 and 2003. The census is measuring quail population response to different management practices. The number of coveys calling from pre-selected points is recorded at daylight. We have 10 points spread evenly across our two 2000 acre study sites. An estimate of quail per acre is calculated after observations are recorded from each point. Site 1 had 0.9 quail per acre in fall 2002, and 0.9 per acre in 2003. Site 2 had 0.7 quail per acre in 2002, and 0.86 per acre in 2003.

Current Activities
Currently conducting the fall 2004 census and preparing to trap wild quail to be radio-tagged and monitored during the 2004-2005 fall/winter period.